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In its function of exporting protein, the rough endoplasmic reticulum works in association with the Golgi apparatus and its vesicles. The smooth endoplasmic reticulum has the separate function of being responsible for lipid production. Also present are a number of filaments of various thickness which together form the cytoskeleton (Chapter 4). One group of these, the microtubules, are 25nm in diameter and are attached to another organelle, the centriole. The centriole functions in the process of cell division which is described in Chapter 8.

L-forms may arise spontaneously by continuously cultivating cells on rich medium (this was how L-forms were first discovered, with the bacterium Streptobacillus moniliformis). More frequently, L-forms can be selected for by cultivating bacteria in an osmotically buffered medium containing penicillin. Penicillin is known to inhibit cell wall (peptidoglycan) synthesis. Usually, if the penicillin is removed soon after L-forms are first formed, then the cells rapidly revert to their normal shape. If, however, the L-forms continue to be cultivated for a long time in the presence of penicillin, the cells may continue to grow in the L-form even when they are transfered to penicillin-free medium (ie the L-forms are stable).

Bacteria have no such connections and it seems reasonable to conclude that the very thick wall of the bacterial cell, while offering strength and protection, may have sentenced them to a life of solitude. The organisation of eukaryotic cells 55 differentiation The matter is not however as simple as this. Let us consider some of the problems posed by multicellularity. In multicellular organisms we find different parts of each organism carry out quite different functions. Think about your own heart, lungs and kidneys or the roots, leaves and flowers of plants.

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